The Villanueva women have a tradition for Mother's Day and Jane is excited to get to participate in her first one as a mom. However, their plans are thwarted when Petra invites everyone to a brunch instead. Jane's TA position is put into jeopardy when a mysterious ad appears in the college newspaper. Rogelio is hurt when Dina doesn't invite him to her birthday party, so he asks Jane to help in an unusual way. Rafael finds himself in big trouble thanks to his half-brother. Meanwhile, Xo finds herself in a compromising position.
Jane has made big plans to celebrate Mateo's first birthday, but things start to fall apart when she learns there might be an issue with her wedding venue. Rogelio faces a challenge on set that could jeopardize Jane's big day. Jane is excited when Alba gives her a lead on a book publisher, but soon begins having doubts about her writing. Jane and Petra try to help Rafael figure out his troubles, but they have differing opinions on how to handle the situation. Meanwhile, Michael is convinced that Derek is really in cahoots with Mutter and has a plan to prove it.
After moving up the wedding day, Jane and Michael find all their carefully made plans thrown into chaos. To make matters worse, Jane is unhappy with her current graduate thesis, but changing it might put a wrinkle in her big day. Rafael still has feelings for Jane, but is torn about whether he should speak now or forever hold his peace. Petra has her own family drama when Anezka has a medical emergency and ends up in the hospital. Meanwhile, Rogelio and Xo are finally coming to terms with the state of their relationship until Xo gets surprising news.
Jane discovers that Michael was shot and finds herself having to make difficult choices. With Rafael focused on keeping Mateo safe, Anezka, who is pretending to be Petra, "helps" the cops with their investigation. However, it isn't long before everyone starts to notice that "Petra" is acting odd. Meanwhile, Rogelio is trying to keep everything out of the press for Jane, but in order to do so finds himself in an awkward position and asks for Xo's help.
When Rafael finally admits that he is over Jane, their co-parenting style is tested when they argue over what the best preschool is for Mateo. Xo is paranoid about Alba finding out her secret and what it will do to their relationship. Rogelio is ready to cross over to American TV, especially after he learns his nemesis is making the transition. Meanwhile, "Anezka" (Yael Grobglas) is trying to find new dirt on Rafael when she begins an unexpected romance.
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